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CHECKLEY, DAVID

Professor

Director, CIMEC

Degrees:

• B.S., University of Washington
• M.S., UCSD, Scripps Institution of Oceanography
• Ph.D., UCSD, Scripps Institution of Oceanography

Research Interests:

• Long-term change in pelagic zooplankton and fish, including climate-biota interactions
• Role of zooplankton in the marine nitrogen cycle
• Effects of weather and air-sea interactions on plankton and fishes
• Ecology of marine zooplankton and fish
• The biological pump, including the role of particles and plankton
• Ocean acidification
• Instrument development

Courses Recently Taught:

• SIO 270A: Fisheries Oceanography
• SIO 279: Sp Top/Biological Oceanography
• SIO 280: Biological Oceanography
• SIO270: Pelagic Ecology
• SIO270A: Fisheries Oceanography
• SIO278: Seminar in Ocean Biosciences
• SIO279: Sp Top/Biological Oceanography
• SIO280: Biological Oceanography

Honors and Awards:

• UCSD Innovator of the Month, Technology Transfer, 2002
• Tech Award Laureate, The Tech Museum, 2002
• NATO Postdoctoral fellow in science, Marine Laboratory, Aberdeen, Scotland, 1978-1979
• Woods Hole Oceanographic Institute Summer Student Fellow, 1968

Mentoring:

• Asch, Sarah (Graduate; 2006-)
• Davison, Peter (Graduate; 2004-)
• Takahashi, Motomitsu (Post Doc; 2004-2006)
• Rykaczewski, Ryan (Graduate; 2003-)
• Glaser, Sarah (Graduate; 2002-)
• Gonzalez-Quiros, Rafael (Post Doc; 2002-2004)
• Rebstock, Ginger (Post Doc; 1999-2000)
• Curtis, K. Alexandra (Graduate; 1997-2003)
• Johnson, Catherine (Graduate; 1996-2003)
• Osgood, Kenric (Post Doc; 1996-1999)
• Rebstock, Ginger (Graduate; 1994-1999)
• Cass-Calay, Shannon (Graduate; 1993-1999)

1. DeVries, A.L., D.M. Checkley, Jr. and J.A. Raymond. Physiology and biochemistry of freezing resistance in Antarctic fishes. Antarctic Jour., 7: 78-79. (1972) RESEARCH ARTICLE 2. Checkley, D.M., Jr. The egg production of a marine, planktonic copepod in relation to its food supply. Laboratory studies. Limnol. Oceanogr., 25: 430-446. (1980) RESEARCH ARTICLE 3. Checkley, D.M., Jr. Food limitation of egg production of a marine, planktonic copepod in the sea off Southern California. Limnol. Oceanogr., 25: 991-998. (1980) RESEARCH ARTICLE 4. Checkley, D.M., Jr. Selective feeding by Atlantic herring. Clupea harengus, larvae on zooplankton in natural assemblages. Mar. Ecol. Prog. Ser., 9: 245-253. (1982) RESEARCH ARTICLE 5. Checkley, D.M., Jr. Relation of growth to ingestion for larvae of Atlantic herring Clupea harengus, and other fish. Mar. Ecol. Prog. Ser., 18: 215-224. (1984) RESEARCH ARTICLE 6. Checkley, D.M., Jr. Nitrogen limitation of zooplankton production and its effect on the marine nitrogen cycle. Arch. Hydrobiol. Beih. Ergebn. Limnol., 21: 103-113. (1985) RESEARCH ARTICLE 7. Checkley, D.M., Jr. and L.C. Entzeroth. Elemental a isotopic fractionations of carbon and nitrogen by marine, planktonic copepods and implications to the marine nitrogen cycle. J. Plank. Res., 7: 553-568. (1985) RESEARCH ARTICLE 8. Angel, M.V., D.M. Checkley, Jr. and S.I. Heany. Chapter one. Plankton migrations introduction. Contr. Mar. Sci. Suppl., 27: 43-44. (1985) BOOK CHAPTER 9. Bird, J.L., D.T. Eppler and D.M. Checkley, Jr. Comparisons of herring otoliths using Fourier series shape analysis. Can. J. Fish. Aq. Sci., 43: 1228-1234. (1986) RESEARCH ARTICLE 10. Checkley, D.M., Jr., S. Raman, G.L. Maillet and K.M. Mason. Winter storm effects on the spawning and larval drift of a pelagic fish. Nature, 335: 346-348. (1988) RESEARCH ARTICLE 11. Checkley, D.M., Jr. and C.A. Miller. Nitrogen isotope fractionation by oceanic zooplankton. Deep-Sea Res., 36: 1449-1456. (1988) RESEARCH ARTICLE 12. Hoss, D.E., D.M. Checkley, Jr. and L.R. Settle. Diurnal buoyancy changes in larval Atlantic menhaden (Brevoortia tyrannus). J. Cons. Int. Explor. Mer., 191: 105-111. (1989) RESEARCH ARTICLE 13. Maillet, G.L. and D.M. Checkley, Jr. Effects of starvation on the frequency and width of growth increments in sagittae of laboratory-reared Atlantic menhaden (Brevoortia tyrannus) larvae. Fish. Bull., 88: 155-165. (1989) RESEARCH ARTICLE 14. Checkley, D.M., Jr. S. Uye, M.J. Dagg, M.M. Mullin, M. Omori, T. Onbe and M.-Y. Zhu. Diel variation of the zooplankton and its environment at neritic stations in the Inland Sea of Japan and the north-west Gulf of Mexico. J. Plank. Res., 14: 1-40. (1992) RESEARCH ARTICLE

15. Maillet, G.L. and D.M. Checkley, Jr. Storm-related variation in the growth rate of otoliths of larval Atlantic menhaden Brevoortia tyrannus: a time series analysis of biological and physical variables and implications for larva growth and mortality. Mar. Ecol. Prog. Ser., 79: 1-16. (1991) RESEARCH ARTICLE 16. Checkley, D.M., Jr., M.J. Dagg and S. Uye. Feeding, excretion and egg production by individuals and populations of the marine, planktonic copepods, Acartia spp. and Centropages furcatus. J. Plank. Res., 14: 71-96. (1992) RESEARCH ARTICLE

17. Checkley, D.M., Jr. Review of K.H. Mann and J.R.N. Lazier, 1991, Dynamics ofd Marine Ecosystems: Biological-Physical Interactions in the oceans, Blackwell. Bull. Amer. Meteorol. Soc., 73: 499-501. (1992) BOOK REVIEW

18. Osgood, K.E., and D.M. Checkley, Jr. Concentration of Calanus pacificus in the Santa Barbara Basin. EOS, 76, p 36. (1996) ABSTRACT 19. Checkley, D.M., Jr., T. Cooper, and C. Lennert. Plankton pattern within and below the surface mixed layer. EOS, 76, 198. (1996) ABSTRACT 20. Osgood, K.E., and D.M. Checkley, Jr. Observations of a deep aggregation of Calanus pacificus in the Santa Barbara Basin. Limnol. & Oceanogr., 42:997-1001. (1996) RESEARCH ARTICLE

21. Osgood, K.E., and D.M. Checkley, Jr. Seasonal variations in a deep aggregation of Calanus pacificus in the Santa Barbara Basin. Mar. Ecol. Prog. Ser., 148:59-69. (1997) RESEARCH ARTICLE 22. Checkley, D.M., Jr., P.B. Ortner, L.R. Settle, and S.R. Cummings. A continuous, underway fish egg sampler. Fish. Oceanogr., 6:58-73. (1997) RESEARCH ARTICLE 23. Cambalik, J.J., D.M. Checkley, Jr., and D. Kamykowski. A new method to measure the terminal velocity of small particles: a demonstration using the ascending eggs of the Atlantic menhaden (Brevoortia tyrannus). Limnol. & Oceanogr. 43:1722-1727. (1998) RESEARCH ARTICLE 24. van der Lingen, C.D., D.M. Checkley, Jr., M. Barange, L. Hutchings, and K. Osgood. Assessing the abundance and distribution of eggs of sardine, Sardinops sagax, and round herring , Etrumeus whiteheadi, on the western Agulhas Bank, South Africa, using a continuous, underway fish egg sampler. Fish. Oceanogr., 7:35-47. (1998) RESEARCH ARTICLE 25. Iwamoto, I., M. M. Trivedi, D.M. Checkley, Jr. Real-time detection and classification of objects in flowing water. Proc. Machine Vision Systems for Inspection and Metrology Conf., SPIE, 3521:214-220. (1998) RESEARCH ARTICLE

26. Checkley, D.M., Jr., P.B. Ortner, F.E. Werner, L.R. Settle, and S.R. Cummings. Spawning habitat of the Atlantic menhaden in Onslow Bay, North Carolina. Fish. Oceanogr. 8:22-36. (1999) RESEARCH ARTICLE 27. Checkley, D.M., Jr., R.C. Dotson, and D.A. Griffith. Continuous underway sampling of eggs of Pacific sardine (Sardinops sagax) and northern anchovy (Engraulis mordax) in spring 1996 and 1997 off southern and central California. Deep-Sea Res. Part II 47:1139-1155. (2000) RESEARCH ARTICLE 28. Checkley, D.M., Jr. 2000. Michael M. Mullin: A biography. PICES Press 8(1):13-17 POPULAR ARTICLE 29. Checkley, D.M., Jr., J.R. Hunter, L. Motos, and C.D. van der Lingen (eds.). 2000. Small Pelagic Fishes and Climate Change Program. Report of a workshop on the use of the Continuous, Underway Fish Egg Sampler (CUFES) for mapping spawning habitats of pelagic fish. GLOBEC Report 14, 1-68 pp. WORKSHOP REPORT (UNREFEREED) 30. Iwamoto, S., D.M. Checkley, Jr., and M.M. Trivedi. 2001. REFLICS: Real-time Flow Imaging and Classification System. Machine Vision and Applications Journal 13(1):1-13. RESEARCH ARTICLE 31. Mullin, M.M., D.M. Checkley, Jr., and M.P. Thimgan. 2003. Temporal and spatial variation in the sizes of California current macrozooplankton: analysis by optical plankton counter. Progress in Oceanography 57(3-4):299-316. RESEARCH ARTICLE 32. Jahncke, J., D.M. Checkley, Jr., and G.L. Hunt, Jr. 2004. Trends in carbon flux to seabirds in the Peruvian upwelling system: effects of wind and fisheries on population regulation. Fisheries Oceanography 13(3):208-223. RESEARCH ARTICLE 33. Johnson, C.L., and D.M. Checkley, Jr. 2004. Vertical distribution of diapausing Calanus pacificus (Copepoda) and implications for transport in the California Undercurrent. Progress in Oceanography 62(1):1-13. RESEARCH ARTICLE 34. Gonzalez-Quiros, R., and D.M. Checkley, Jr. 2006. Occurrence of fragile particles inferred from optical plankton counters used in situ and to analyze net samples collected simultaneously. Journal of Geophysical Research. III, d.o.i:10.1029/2005JC003084. RESEARCH ARTICLE 35. Curtis, K.A., D.M. Checkley, Jr., and P. Pepin. 2007. Predicting the vertical profiles of anchovy (Engraulis mordax) and sardine (Sardinops sagax) eggs in the California Current System. Fisheries Oceanography 16:68-84. RESEARCH ARTICLE 36. Reiss, C., D.M. Checkley, Jr., and S. Bograd. 2008. Remotely-sensed spawning habitat of Pacific sardine (Sardinops sagax) and Northern anchovy (Engraulis mordax) within the California Current. Fisheries Oceanography 17:126-136. RESEARCH ARTICLE 37. Rykaczewski, R.R., and D.M. Checkley, Jr. 2008. Influence of ocean winds on the pelagic ecosystem in upwelling regions. Proc. Nat. Acad. Sci. 105:1965-1970. RESEARCH ARTICLE 38. Checkley, D.M., Jr., J. Alheit, Y. Oozeki, and C. Roy (eds.) In press. Climate Change and Small Pelagic Fish, Cambridge University. Press (Checkley lead author of Preface and Chapters 3 (Habitats) and 15 (Synthesis)); 82 authors total; ca. 450 pages. RESEARCH ARTICLE